After escaping their cell, Skipper and Julien retreat to the surface where they find the rats hugging and kissing the zoo animals. The real Red is then captured. There, Private soon becomes smitten with Shawna, a new intern who tends to Private's injuries. The plan succeeds, as does Marlene's original goal of bringing the zoo together. Left alone in an alley, Private is passed a cassette player by the other penguins containing audio instructions on how to outwit this enemy. Throughout, only Private believes such is wrong. His mother soon locates him there, and tells the penguins that Eggy has been behaving that way ever since their egg-watching adventure.
Soon, the nature of Private and the Amarillo Kid's past is shown: Private was once a skilled mini golf player, but quit the sport after a tough game against the Amarillo Kid forced him to have to ricochet a ball off a young opossum girl's ice cream cone, ruining it. As they surround him wishing to do him harm, the Love-u-Laser's effects are finally reversed when it falls into the fountain and causes a loathe-reversing beam to be cast across the zoo. But in the process of doing so, the Love-u-Laser jams and then fires unexpectedly at Private, who within moments angrily tackles and yells at Kowalski. Soon, the team is greeted by friends and enemies Lulu, Hans, Savio, Clemson, and Rhonda who cheerfully welcome the penguins to Hoboken and tell them about Frances, a new zookeeper who has made their lives enjoyable, including by letting them use massage chairs. Whilst attempting to do so at the photographer's house, however, Kowalski inadvertently sends an enlarged version of the incriminating area of the photo to the ad executives. When Skipper and Julien bicker about each other's unfitness at a zoovenir shop meeting, Mason introduces a motion that the two change places for a day to gain better mutual understanding. Despite their best attempts, the penguins are unable to procure a voice chip but Private obtains a different toy with a similar chip, so Kowalski transplants that chip into Rico's doll.
Private desires a promotion after Skipper forbids him to look at a joke contained in a classified mission file. Meanwhile, Julien, Maurice, and all the animals are in the shop itching like crazy. Objects once invisible, including the ray itself, become visible again. When Roy soon awakens and begins to charge again, however, it is the penguins who have the last laugh when a stuffed penguin takes the blow and Julien momentarily believes it was Skipper. When the time capsule is opened in the present day, the race is soon on between the penguins, lemurs, Marlene, and Fred to reach the legendary treasure before King Rat and his minions do. But since fish are brain food, when Kowalski starts eating them, he begins to regrow his normal brain capacity.
Blowhole, was abandoned after the penguins discovered that dolphins do not use toilets. Julien, furious from catering Maurice's every need, proceeds to chase him away, throwing walnuts at him. After Julien gains possession of it and inadvertently stops and then restarts time, Kowalski attempts to get the stopwatch back, but the timepiece ultimately winds up irreparably broken, permanently trapping the two in a moment of time. But when Private saves Hunter's father from falling off a ledge in the process, he accepts that natural instincts are not everything. Not wanting Hunter to have to travel to Antarctica alone, Private launches with her, forcing the others to have to follow by submarine.
Left alone in an alley, Private is passed a cassette player by the other penguins containing audio instructions on how to outwit this enemy. At the lemur habitat, Julien complains to Rico about a nearby billboard which he finds bothersome, which Rico soon destroys with dynamite. The penguins soon catch and they attempt to stop the car from hurting Rico. He tells Maurice to be Rico, Mort to be Skipper, Fred the squirrel to be Kowalski, and he made himself Private. But Mort accidentally blows his cover and gets caught. This article's plot summary may be.
Wishing for gumballs, Mort tosses the penny in and, to the serendipitous surprise of Private, gets them, leading Private to conclude that the fountain is magic. Their version of play involves placing the two through uncomfortable scenarios for their amusement, including trying to make them fight each other, which Skipper refuses to engage in but Julien goes along with because of his fascination with the shiny objects the twins reward him with. Once Savio arrives, however, he gets past the animals' defenses and it is revealed that Julien had confused Savio with Fred. The penguins then chase after Frankie, with Skipper flying through the air using a jet-pack, but the pigeon soon ends up releasing his blueberry-infused vendetta. The Penguins of Madagascar: Episode Info.
Through flashbacks beginning when Skipper discovered that Kowalski still possessed Jiggles, it is shown that Kowalski believed Jiggles could now be safely kept because he had altered it to consume whipped cream instead of fruit. In the end, Skipper and the others learn their lesson and agree to respect Private's wishes and stop using the hyper-cute. Having just made a wish on a star, Julien believes the necklace is his gift from above. After Julien gains possession of it and inadvertently stops and then restarts time, Kowalski attempts to get the stopwatch back, but the timepiece ultimately winds up irreparably broken, permanently trapping the two in a moment of time. After feeling sad that she is not understood, Hunter, along with Private and Julien, who had been a submarine stowaway, rescue the other penguins, though the hungry seals pursue. The zoo animals then turn on the rats, causing them to retreat back into the sewer.
When a boa constrictor arrives at the zoo from Hoboken, Skipper and Kowalski warn of the snake's dangers at a zoovenir shop meeting. There, the team is quickly noticed by humans, boxed, and brought to the feared Hoboken Zoo. Soon after locating him, however, their situation becomes more difficult when Leonard falls asleep in the daylight. The penguins set off on a sailing trip to the Massachusetts coast, but poor weather abruptly tosses them onto the streets of Hoboken. Though initially frustrated by what Julien had caused, Kowalski soon uses the sudden eternity to partake in buddy activities with Julien and create 7,009 inventions. Meanwhile, the penguins observe the boy, Ronald, telling his teacher about having seen their operations, but the teacher rejects his account.